Greg RaglandGreg Ragland
Assistant Professor
Department of Integrative Biology
University of Colorado, Denver
Campus Box 171
Denver, CO 80217-3364
PHONE: (303) 556-5970
gregory DOT ragland AT ucdenver DOT edu

Greg Ragland. I have been an assistant professor in the Department of Integrative Biology at CU Denver since 2016. When I am not immersed in bugs and data, I am usually wrangling my daughters, hiking, running, or cycling.



Eddy Dowle is studying local adaptation and population structure in the Mountain Pine Beetle and developmental transcriptomics of diapause regulation in apple maggot flies. She has also worked on the phylogeography of New Zealand stick insects and New Caledonian snails.


Visiting Scholars

You-Jin Hao, Chongqing Normal University , Chongqing, China. You-Jin is an insect molecular biologist who studies mechanisms of seasonality and diapause regulation. He is currently developing RNAi approaches in Rhagoletis pomonella and is collaborating on studies of gene expression and developmental dynamics during diapause development.


Graduate Students


Phil Freda (currently based at KSU). (website) I am broadly interested in the evolution of complex life cycles and levels of constraint (or lack thereof) across metamorphic boundaries. In other words, is performance under a given set of environmental conditions constrained across metamorphosis or is there the potential for adaptive decoupling in distinct life stages? I aim to answer this question by estimating genetic correlations in thermal limits across metamorphosis using the fully sequenced Drosophila Genetic Reference Panel (DGRP) of Drosophila melanogaster.

McCall Calvert (Mac). — MS student.

Marianne Blackburn. — MS student.


Nooshin Sanaei studies patterns of gene expression and cell proliferation associated with cell cycle arrest during insect dormancy.


Lab Alumni


Adam Schieferecke was a summer research assistant who mined genomic and transcriptomic data for diapause candidate genes and worked with Phil on thermal physiology experiments in spotted wing drosophila (Drosophila suzukii). He is now completing a BS/MS at KSU after earning a prestigious Barry M Goldwater scholarship.

Mariah Brown

Mariah Brown worked on various thermal ecology experiments in D. melanogaster and suzuki, and  continues to work towards her BS at KSU.

Evan Keep worked on gene orthology and comparative transcriptomics as a summer REU student. He has recently completed his BS and is now employed at LabAnswer.



Dan Hahn and Tom Powell (U. Florida), Jeff Feder and Mike Pfrender (Notre Dame), Barbara Bentz (USDA), Karen Mock (Utah State U.), Stewart Berlocher and Hugh Robertson (U. Illinois), Christian Stauffer (BOKU, Vienna). Photo credit: Hannes Schuler.